Publications: Scott J. Steppan

Publications are listed below with links to futher information on them.


Schenk, J.J., Rowe, K.C., and Steppan, S.J. Ecological opportunity and incumbency in the diversification of repeated continental colonizations by muroid rodents. Systematic Biology. In press.
pre-press early access pdf (2 Mb)


Ortiz, P. E., Jayat, J. P., and Steppan, S. J. A new fossil Phyllotini (Rodentia: Sigmodontinae) from Late Pliocene in the Andes of northwestern Argentina. Journal of Vertebrate Paleontology. 32 (6): 1429-1441.
PDF (1.7 Mb)

2012 G. M. Erickson, P. M. Gignac, S. J. Steppan, A. K. Lappin, K. A. Vliet, J. D. Brueggen, B. D. Inouye, D. Kledzik, G. J. W. Webb. Insights into the ecology and evolutionary success of crocodilians revealed through bite force and tooth pressure experimentation PLoS ONE 7(3): e31781. doi:10.1371/journal.pone.0031781.
PDF (570 kb)

Steppan, S..J., Kenagy, G. J., Zawadzki, C., Robles, R., Lyupanova, E.A., and Hoffmann, R.S. Molecular data resolve placement of the Olympic marmot and estimate dates of trans-Beringian interchange. Journal of Mammalogy. 92(5):1028-1037.
PDF (1.3 Mb)

2011 Burns, J. H., Faden, R. B., and Steppan, S. J. Phylogeny of Commelinaceae inferred from nuclear ribosomal and chloroplast DNA sequences. Systematic Botany. 36(2):268-276..
PDF (560 kb)

Lawrence R. Heaney, Danilo S. Balete, Eric A. Rickart, Phillip A. Alviola, Mariano Roy M.  Duya, Melizar V. Duya, M. Josefa Veluz, Lawren VandeVrede, and Scott J. Steppan. Seven new species and a new subgenus of forest mice (Rodentia: Muridae: Apomys) from Luzon Island, Philippines. Fieldiana: Life and Earth Sciences 2: 1–60.
PDF (8.5 Mb)
FSU Radio news story and interview (3.2 Mb mp3)
FSU video interview (link)

2010 Cooper, J., and Steppan, S. J. Developmental constraint on the evolution of marsupial forelimb morphology. Australian Journal of Zoology. 58: 1-15.
PDF (444 kb)
2010 van der Linde, K., Houle, D., Spicer, G., and Steppan, S. J.  A supermatrix analysis of the family Drosophilidae. Genetics Research. 92: 25-38.
PDF (210 kb)
2008 Harshman, J., Braun, E. L., Braun, M. J., Huddleston, C. J., Bowie. R. C. K., Chojnowski, J. L., Hackett, S. J., Han, K-L., Kimball, R. T, Marks, B. D., Miglia, K. J., Moore, W. S., Reddy, S., Sheldon, F. H., Steadman, D. W., Steppan, S. J., Witt, C. J., and Yuri, T. Phylogenomic evidence for multiple losses of flight in ratite birds. Proceedings of the National Academy of Science 105 (36): 13462-13467.
PDF (680 kb)
Supporting Information (3.6 Mb pdf)
2008 Steppan, S. J. Aligning the spaces: a comment on Polly - Developmental dynamics and G-matrices. Evolutionary Biology. 35(2):108-110.
PDF (100 kb)

Good, J. M., Hird, S., Demboski, J. R., Steppan, S. J., Martin-Nims, T.R., and Sullivan, J. Ancient hybridization and mitochondrial capture between two distantly related species of chipmunks (Tamias: Rodentia). Molecular Ecology. 17(5):1313-1327.
PDF (450 kb)

2008 Rowe, K., Reno, M. L., Richmond, D. M., Adkins, R. M., and Steppan, S. J. Pliocene colonization and adaptive radiations in Australia and New Guinea (Sahul): Multi-locus systematics of the Old Endemic rodents (Muroidea: Murinae). Molecular Phylogenetics and Evolution. 47:84-101.
PDF (630 kb)
2007 Steppan, S. J., Ramirez, O., Banbury, J., Huchon, D., Pacheco, V., Walker, L, and Spotorno, A.O. A molecular reappraisal of the systematics of the leaf-eared mice Phyllotis and their relatives. Pp. xx-xx in The quintessential naturalist: honoring the life and legacy of Oliver P. Pearson (Kelt, D. A., E. Lessa, J. A. Salazar-Bravo, and J. L. Patton, eds.).  University of California Publications in Zoology.
PDF (580 kb)
in press Steppan, S.J. and Ramirez-Baca, O. Genus Phyllotis. In South American Mammals. J. L. Patton ed..

2005 Steppan, S. J., Adkins, R. M., Spinks, P. Q., and Hale, C. Multigene phylogeny of the Old World mice Murinae reveals distinct geographic lineages and the declining utility of mitochondrial genes compared to nuclear genes. Molecular Phylogenetics and Evolution. 37:370-388.
PDF(1.3 Mb)
2004 Steppan, S. J., Seeing the Forest for the Trees. Book review of "Assembling the Tree of Life." Science. 307:677-678.
PDF (250 kb)
2004 Steppan, S. J., Adkins, R. M., and Anderson, J. Phylogeny and divergence-date estimates of rapid radiations in muroid rodents based on multiple nuclear genes. Systematic Biology. 53(4):533-553.
PDF (300 kb)
Cover illustration (jpg)
2004 Holliday, J. A. and Steppan, S. J. Evolution of hypercarnivory: the effect of specialization on morphological and taxonomic diversity. Paleobiology. 30: 108-128
PDF (444 kb).
2004 Steppan, S. J. Phylogenetic comparative analysis of multivariate data. In Phenotypic Integration. (eds., Piggliuci, M. and Preston, K. A.). Oxford Univ. Press.
Manuscript (500 kb pdf)
2004 Steppan, S. J., Storz, B. L., and Hoffmann, R. S. Nuclear DNA phylogeny of the squirrels (Mammalia: Rodentia) and the evolution of arboreality from c-myc and RAG1. Molecular Phylogenetics and Evolution. 30: 703-719
pdf (420 kb)
2003 Steppan, S. J., Zawadzki, C., and Heaney, L. R. Molecular phylogeny of the endemic Philippine rodent Apomys (Muridae) and the dynamics of diversification in an oceanic archipelago. Biological Journal of the Linnean Society 80:699-715.
PDF (310 kb)
2002 Steppan, S.J., Houle, D., Phillips, P.C. Comparative quantitative genetics: evolution of the G matrix. Trends in Ecology and Evolution 17:320-327.
pdf (80 kb)
2002 Kuch, M. Betancourt, J., Lattore, C., Steppan, S.J., Poinar, H.N. Molecular Analysis of a 11,700-yr old rodent midden from the Atacama Desert Chile. Molecular Ecology 11: 913-924 .
pdf (1 Mb)
2000 Steppan, S.J. and Sullivan, J. The emerging statistical perspective in systematics and the status of Andalgalomys (Rodentia: Sigmodontinae): a reply to Mares and Braun. Journal of Mammalogy.
pdf (860 kb)
2000 Ortiz, P.E., Pardiñas, U.F.J., and Steppan, S.J. A new extinct phyllotine (Rodentia: Muridae) from the Pleistocene of Argentina and a consideration of the phylogenetic relationships of the "Reithrodon group." Journal of Mammalogy. 81(1): 37-15.
pdf (1Mb)
2000 Steppan, S.J.  Flexural stiffness patterns of butterfly wings. Journal of Research on the Lepidoptera.
pdf (484 kb)
1999 Steppan, S.J., Hakhverdyan, M, Lyapunova, E., Vorontsov, N., Fraser, D.G., Hoffmann, R.S., and Braun, M.J. Molecular phylogeny of the marmots (Rodentia: Sciuridae): tests of evolutionary and biogeographic hypotheses. Systematic Biology.
Abstract and Figures (html)
Full text (pdf)
1998 Steppan, S. J. and Pardiñas, U. F.    Two new fossil murids from the Pleistocene of Argentina: phylogeny and paleoecology. Journal of Vertebrate Paleontology. 18(3):640-649.
Abstract and Figures
1998 Steppan, S.J.   Phylogenetic relationships and species limits within Phyllotis (Rodentia: Sigmodontinae): concordance between mtDNA sequence and morphology. Journal of Mammalogy. 79(2):573-593.
Abstract, Figures, and Emended Diagnosis
1997 Steppan, S.J.    Phylogenetic analysis of phenotypic covariance structure. I. Contrasting results from matrix correlation and Common Principal Component analyses. Evolution. 51(2):571-586.
Full text, hi-res (3.1 Mb pdf); low-res (900 kb pdf)
1997 Steppan, S.J.   Phylogenetic analysis of phenotypic covariance structure. II. Reconstructing matrix evolution. Evolution. 51(2):587-594.
Full text, hi-res (1.6 Mb pdf); low-res (450 kb pdf)
1996 Steppan, S.J.   A new species of Holochilus (Rodentia: Sigmodontinae) from the middle Pleistocene of Bolivia and its phylogenetic significance. Journal of Vertebrate Paleontology. 16(3):522-530.
Abstract and Figures
1995 Steppan, S.J.  Revision of the tribe Phyllotini (Rodentia: Sigmodontinae) with a phylogenetic hypothesis for the Sigmodontinae. Fieldiana: Zoology. n.s. 80:1-112.
Abstract and Figures; full pdf (8.5 Mb); or view pages on Biodiversity Heritage Library
1995 Ph.D. dissertation. University of Chicago. Phylogenetic relationships of the phyllotine rodents (Sigmodontinae) and the evolution of phenotypic patterns of covariation in Phyllotis.
1993 Steppan, S.J.   Phylogenetic relationships among the Phyllotini (Rodentia: Sigmodontinae) using morphological characters. Journal of Mammalian Evolution. 1(3):187-213.
1991 Steppan, S.J.    Geographic distribution of flower morphological traits in subspecies of Lotus scoparius. Journal of Biogeography. 18:321-331.
Abstract and Figures