Publications
In press

Steppan, S.J. and Ramirez-Baca, O. Genus Phyllotis. In South American Mammals. J. L. Patton ed..

Published

40. G. M. Erickson, P. M. Gignac, S. J. Steppan, A. K. Lappin, K. A. Vliet, J. D. Brueggen, B. D. Inouye, D. Kledzik, G. J. W. Webb. Insights into the ecology and evolutionary success of crocodilians revealed through bite force and tooth pressure experimentation PLoS ONE 7(3): e31781. doi:10.1371/journal.pone.0031781.

39. Ortiz, P. E., Jayat, J. P., and Steppan, S. J. A new fossil Phyllotini (Rodentia: Sigmodontinae) from Late Pliocene in the Andes of northwestern Argentina. Journal of Vertebrate Paleontology


38. Steppan, S..J., Kenagy, G. J., Zawadzki, C., Robles, R., Lyupanova, E.A., and Hoffmann, R.S. Molecular data resolve placement of the Olympic marmot and estimate dates of trans-Beringian interchange. Journal of Mammalogy 92(5): 1028-1037.

37. Burns, J. H., Faden, R. B., and Steppan, S. J. Phylogeny of Commelinaceae inferred from nuclear ribosomal and chloroplast DNA sequences. Systematic Botany 36(2): 268-276.

36. Lawrence R. Heaney, Danilo S. Balete, Eric A. Rickart, Phillip A. Alviola, Mariano Roy M. Duya, Melizar V. Duya, M. Josefa Veluz, Lawren VandeVrede, and Scott J. Steppan. Seven new species and a new subgenus of forest mice (Rodentia: Muridae: Apomys) from Luzon Island, Philippines. Fieldiana: Life and Earth Sciences. 2: 1-60.

35. Swain, TD. Evolutionary transitions in symbioses: dramatic reductions in bathymetric and geographic ranges of Zoanthidea coincide with loss of symbioses with invertebrates. Molecular Ecology.


34. Cooper, J., and Steppan, S. J. Developmental constraint on the evolution of marsupial forelimb morphology. Australian Journal of Zoology. 58: 1-15.

33. Holliday, J. A. Evolution in Carnivora: Identifying a morphological bias. In Carnivoran Evolution: New Views on Phylogeny, Form, and Function (A. Goswami and A. Friscia, eds.). Cambridge: Cambridge University Press, pp. 141-164.

32. van der Linde, K., Houle, D., Spicer, G., and Steppan, S. J. A supermatrix analysis of the family Drosophilidae. Genetics Research. 92: 25-38


32. Swain T. D. Phylogeny-based species delimitations and the evolution of host associations in symbiotic zoanthids (Anthozoa, Zoanthidea) of the wider Caribbean region Zoological Journal of the Linnean Society. 156: 223-238.


31. Harshman, J., Braun, E. L., Braun, M. J., Huddleston, C. J., Bowie. R. C. K., Chojnowski, J. L., Hackett, S. J., Han, K-L., Kimball, R. T, Marks, B. D., Miglia, K. J., Moore, W. S., Reddy, S., Sheldon, F. H., Steadman, D. W., Steppan, S. J., Witt, C. J., and Yuri, T. Phylogenomic evidence for multiple losses of flight in ratite birds. Proceedings of the National Academy of Science. 105 (36): 13462-13467.

30. Steppan, S. J. Aligning the spaces: a comment on Polly - Developmental dynamics and G-matrices. Evolutionary Biology. 35(2): 108-110.

29. Good, J. M., Hird, S., Demboski, J. R., Steppan, S. J., Martin-Nims, T.R., and Sullivan, J. Ancient hybridization and mitochondrial capture between two distantly related species of chipmunks (Tamias: Rodentia). Molecular Ecology. 17(5): 1313-1327.

28. Rowe, K., Reno, M. L., Richmond, D. M., Adkins, R. M., and Steppan, S. J. Pliocene colonization and adaptive radiations in Australia and New Guinea (Sahul): Multi-locus systematics of the Old Endemic rodents (Muroidea: Murinae). Molecular Phylogenetics and Evolution. 47: 84-101.


27. Steppan, S. J., Ramirez, O., Banbury, J., Huchon, D., Pacheco, V., Walker, L, and Spotorno, A.O. A molecular reappraisal of the systematics of the leaf-eared mice Phyllotis and their relatives. Pp. xx-xx in The quintessential naturalist: honoring the life and legacy of Oliver P. Pearson (Kelt, D. A., E. Lessa, J. A. Salazar-Bravo, and J. L. Patton, eds.). University of California Publications in Zoology.

26. Storz, B. L. and Travis J.  Trait-specific and temporally dissociated developmental program modification in Spea multiplicata tadpoles suggests modularity.  TSW Development and Embryology. 7: 715-726.

25. Swain T. D. and Wulff J. L. Diversity and specificity of Caribbean sponge-zoanthid symbioses: a foundation for understanding the adaptive significance of symbioses and generating hypotheses about higher-order systematics. Biological Journal of the Linnean Society. 92: 695-711.


24. Weisrock, D. W., Shaffer, H. B., Storz, B. L., Storz, S. R., and Voss, S. R. Multiple nuclear gene sequences identify phylogenetic species boundaries in the rapidly radiating clade of Mexican ambystomatid salamanders.  Molecular Ecology. 15(9): 2489-2503. 


23. Steppan, S. J., Adkins, R. M., Spinks, P. Q., and Hale, C. Multigene phylogeny of the Old World mice Murinae reveals distinct geographic lineages and the declining utility of mitochondrial genes compared to nuclear genes. Molecular Phylogenetics and Evolution. 37: 370-388.


22. Storz, B. L.  Reassessment of the environmental mechanisms controlling developmental polyphenism in spadefoot toad tadpoles.  Oecologia. 141(3): 402-410.

21. Steppan, S. J., Seeing the Forest for the Trees. Book review of "Assembling the Tree of Life." Science. 307: 677-678.

20. Steppan, S. J., Adkins, R. M., and Anderson, J. Phylogeny and divergence-date estimates of rapid radiations in muroid rodents based on multiple nuclear genes. Systematic Biology. 53(4):533-553.

19. Holliday, J. A. and Steppan, S. J. Evolution of hypercarnivory: the effect of specialization on morphological and taxonomic diversity. Paleobiology. 30: 108-128.

18. Steppan, S. J. Phylogenetic comparative analysis of multivariate data. In Phenotypic Integration. (eds., Piggliuci, M. and Preston, K. A.). Oxford Univ. Press.

17. Steppan, S. J., Storz, B. L., and Hoffmann, R. S. Nuclear DNA phylogeny of the squirrels (Mammalia: Rodentia) and the evolution of arboreality from c-myc and RAG1. Molecular Phylogenetics and Evolution. 30: 703-719.


16. Steppan, S. J., Zawadzki, C., and Heaney, L. R. Molecular phylogeny of the endemic Philippine rodent Apomys (Muridae) and the dynamics of diversification in an oceanic archipelago. Biological Journal of the Linnean Society. 80: 699-715.


15. Steppan, S.J., Houle, D., Phillips, P.C. Comparative quantitative genetics: evolution of the G matrix. Trends in Ecology and Evolution. 17: 320-327.

14. Kuch, M. Betancourt, J., Lattore, C., Steppan, S.J., Poinar, H.N. Molecular Analysis of a 11,700-yr old rodent midden from the Atacama Desert Chile. Molecular Ecology. 11: 913-924.


13. Steppan, S.J. and Sullivan, J. The emerging statistical perspective in systematics and the status of Andalgalomys (Rodentia: Sigmodontinae): a reply to Mares and Braun. Journal of Mammalogy.

12. Ortiz, P.E., Pardiñas, U.F.J., and Steppan, S.J. A new extinct phyllotine (Rodentia: Muridae) from the Pleistocene of Argentina and a consideration of the phylogenetic relationships of the "Reithrodon group." Journal of Mammalogy. 81(1): 37-15.

11. Steppan, S.J. Flexural stiffness patterns of butterfly wings. Journal of Research on the Lepidoptera.


10. Steppan, S.J., Hakhverdyan, M, Lyapunova, E., Vorontsov, N., Fraser, D.G., Hoffmann, R.S., and Braun, M.J. Molecular phylogeny of the marmots (Rodentia: Sciuridae): tests of evolutionary and biogeographic hypotheses. Systematic Biology.


9. Steppan, S. J. and Pardiñas, U. F. Two new fossil murids from the Pleistocene of Argentina: phylogeny and paleoecology. Journal of Vertebrate Paleontology. 18(3): 640-649.

8. Steppan, S.J. Phylogenetic relationships and species limits within Phyllotis (Rodentia: Sigmodontinae): concordance between mtDNA sequence and morphology. Journal of Mammalogy. 79(2): 573-593.


7. Steppan, S.J. Phylogenetic analysis of phenotypic covariance structure. I. Contrasting results from matrix correlation and Common Principal Component analyses. Evolution. 51(2): 571-586.

6. Steppan, S.J. Phylogenetic analysis of phenotypic covariance structure. II. Reconstructing matrix evolution. Evolution. 51(2): 587-594.


5. Steppan, S.J. A new species of Holochilus (Rodentia: Sigmodontinae) from the middle Pleistocene of Bolivia and its phylogenetic significance. Journal of Vertebrate Paleontology. 16(3): 522-530.


4. Steppan, S.J. Revision of the tribe Phyllotini (Rodentia: Sigmodontinae) with a phylogenetic hypothesis for the Sigmodontinae. Fieldiana: Zoology. n.s. 80: 1-112.

3. Ph.D. dissertation. University of Chicago. Phylogenetic relationships of the phyllotine rodents (Sigmodontinae) and the evolution of phenotypic patterns of covariation in Phyllotis.


2. Steppan, S.J. Phylogenetic relationships among the Phyllotini (Rodentia: Sigmodontinae) using morphological characters. Journal of Mammalian Evolution. 1(3):187-213.


1. Steppan, S.J. Geographic distribution of flower morphological traits in subspecies of Lotus scoparius. Journal of Biogeography. 18: 321-331.

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